Wine – Unable to Install winehq-staging on Ubuntu 16.04 64bit

I have been running wine on Ubuntu 16.04, installed via sudo apt-get install wine, but I would like to switch to wine-staging. I am attempting to follow the instructions at https://wiki.winehq.org/Ubuntu

I first uninstalled wine and made sure all packages were up to date:

sudo apt-get remove wine wine-mono wine-gecko winetricks
sudo apt-get autoremove
sudo apt-get update
sudo apt-get upgrade

I then rebooted and ran the following series of commands:

sudo dpkg --add-architecture i386
wget -nc https://dl.winehq.org/wine-builds/Release.key
sudo apt-key add Release.key
sudo apt-add-repository https://dl.winehq.org/wine-builds/ubuntu/
sudo apt-get update

However, the command to install winehq-staging fails with unmet dependency on wine-staging:

sudo apt-get install --install-recommends winehq-staging

The following packages have unmet dependencies:
 winehq-staging : Depends: wine-staging (= 2.13.0~xenial)
E: Unable to correct problems, you have held broken packages.

Installing wine-staging fails with unmet dependency on wine-staging-i386:

sudo apt-get install wine-staging

The following packages have unmet dependencies:
 wine-staging : Depends: wine-staging-i386 (= 2.13.0~xenial)
E: Unable to correct problems, you have held broken packages.

Installing wine-staging-i386 fails with various unmet dependencies on -i386 packages. Showing a few of them (the list is much longer):

sudo apt-get install wine-staging-i386

The following packages have unmet dependencies:
 wine-staging-i386:i386 : Depends: libasound2:i386 (>= 1.0.16) but it is not installable
                          Depends: libglib2.0-0:i386 (>= 2.12.0) but it is not installable
                          Depends: libgphoto2-6:i386 (>= 2.5.9) but it is not installable
                          Depends: libgphoto2-port12:i386 (>= 2.5.9) but it is not installable

(snip)

                          Recommends: libxslt1.1:i386 (>= 1.1.25) but it is not installable
                          Recommends: libxxf86vm1:i386 but it is not installable
E: Unable to correct problems, you have held broken packages.

If I try to install the first one, it is not available:

sudo apt-get install libasound2:i386

Package libasound2:i386 is not available, but is referred to by another package.
This may mean that the package is missing, has been obsoleted, or
is only available from another source
However the following packages replace it:
  libasound2-data

E: Package 'libasound2:i386' has no installation candidate

However, package libasound2-data is already installed and the latest version.

I'm not sure what to try next.