Trouble adding a printer with lpadmin

I'm trying to add a printer through the Terminal using lpadmin, and it absolutely refuses to work. The printer adds just fine, everything looks perfect from what I can see, but any attempt to print results in a "Printer did not respond" error.

Here's the line I'm using to add the printer:

lpadmin -p _1BIT_16x09 -D 1BIT_16x09 -L "Export" -o printer-is-shared=false -E -v lpd://10.96.99.60/1BIT_16X09 -P ~/PPDs/_1BIT_16x09.ppd

The reason the -p name has a _ in front of it is because if I add it normally, it adds this, I'm assuming because the name starts with a number. I have crafted it to create the printer to appear exactly the same way within the cups web interface and the /etc/cups/printers.conf file.

If I add the printer using the above command (which doesn't work), this is what I get in printers.conf:

<Printer _1BIT_16x09>
UUID urn:uuid:13a13ddb-3110-37b0-7aab-96bae9e30955
Info 1BIT_16x09
Location Export
MakeModel Agfa ApogeeX
DeviceURI lpd://10.96.99.60/1BIT_16X09
State Stopped
StateMessage The printer did not respond.
StateTime 1464903468
ConfigTime 1464903399
Reason paused
Type 8450124
Accepting Yes
Shared No
JobSheets none none
QuotaPeriod 0
PageLimit 0
KLimit 0
OpPolicy default
ErrorPolicy stop-printer
</Printer>

If I add it normally through the GUI (which does work), this is what I get:

<Printer _1BIT_16x09>
UUID urn:uuid:c5565ce1-256d-3d66-4fb8-74fde3242cc5
Info 1BIT_16x09
Location Export
MakeModel Agfa ApogeeX
DeviceURI lpd://10.96.99.60/1BIT_16X09
State Idle
StateTime 1464904755
ConfigTime 1464904755
Type 8450124
Accepting Yes
Shared No
JobSheets none none
QuotaPeriod 0
PageLimit 0
KLimit 0
OpPolicy default
ErrorPolicy stop-printer
</Printer>

As far as I can tell, aside from the UUID they are identical, but one works and the other doesn't. It's driving me bonkers. Any ideas?