MacOS – list all the Bonjour-enabled services that are running

The dns-sd command-line tool can be helpful for this, but learning to use it is a bit tricky.

Running dns-sd -B _services._dns-sd._udp will return a list of all available service types that currently being advertised. (The list is per interface, so there will be some redundancy.) If this is done on a Mac with no active network connection, the list will of course only contain services running on that machine.

Using that list, you can request information about the individual services types by running things like dns-sd -B _home-sharing._tcp (which lists iTunes Home Sharing instances), and then, given an instance name, you can run dns-sd -L "Wes Campaigne’s Library" _home-sharing._tcp to lookup information for a particular instance.

To be honest, though, this whole process is rather tedious, and it's exactly what Discovery (formerly Bonjour Browser) was built to do, so I highly recommend using that. Whichever way you do it, though, it may not always be clear what program is responsible for a given service entry.

You can use the strategy given in binarybob's answer to try to map service entries to running processes based on port number, but this may not always work. Another strategy is to run something like dns-sd -B _home-sharing._tcp which remains open and reports when instances are added or removed, then, one by one, quit various apps and (assuming it's not a system service) see which one's quitting triggers the removal message.