Since the Screen Sharing feature seems to be the only one unable to connect, the best way to diagnose its behavior is by using the application directly.
Please, launch screen sharing located in:
/System/Library/CoreServices/ScreenSharing.app
The application has no visible "UI" so don't expect a fancy Cocoa interface. Instead, once you have loaded it, all you'll see is this "poor" icon in the Dock:
Click on it and then go to Preferences in the Mac menu bar. You should be presented with:
Notice the second option to only encrypt passwords and keystrokes. That's fine, but as you can see, there's no option to skip encryption altogether. Some VNC versions have incompatibilities with encryption, but this would be rather shocking since you're talking about OS X vs OS X.
In any case, once you have the settings like that, try connecting, again, in the menu bar: Connection -> New (or ⌘cmd + N).
A simple dialog appears asking you to enter the Host's name. Following your screenshots, try: 192.168.1.99 and hit enter.
We'll see what happens from here. Also, have the Console.app open on both computers, to see if there are any System messages going on (or anything related).
VNC server changed on lion. You can now log in without a vnc password but instead he os users each get their own loginwindow allowing simultaneous log in distinct from what the local display shows.
Some discussions on what exactly changed can be found here: Are the changes to Lion's screen sharing documented anywhere publicly?
I do know the mac screen sharing client as well as edovia's iOS screens apps work well with the new lion changes, so there's hope other third party clients can get updated to work better with the new changes.
Best Answer
It is none of them, however it is compatible with VNC.